PD Dr. Peter Döbbeler

Researcher

Systematics, Biodiversity and Evolution of Plants

Office address:

Menzinger Str. 67

Room 249

80638 Munich

Publications

2026

Döbbeler, P. (2026). Beyond the moss: an exploration of bryophilous ascomycetes. – Nova Hedwigia 122(1–2): 79–104.

2025

Döbbeler, P. (2025). Bryophilous ascomycetes from southern South America and adjacent (Sub)Antarctic regions. – Herzogia 38: 275–307.

Greiff, G. R. L., Stöckli, E., Bartholomeeusen, F. & Döbbeler, P. (2025). Bryocentria insolens (Hypocreales) – a new bryophilous ascomycete on Lewinskya and a preliminary phylogenetic analysis of European Bryocentria. – Plant and Fungal Systematics 70(1): 28–37.

2024

Baral, H.-O., Krieglsteiner, L.G., Sochorová, Z., Döbbeler, P., Priou, J.-P., Stöckli, E., Rubio, E., Valade, F., Bauer, G., Hairaud, M., Isaksson, R., Greiff, G. Krieglsteiner, K., Sochor, M. & Lendemer, J.C. (2024). Luteodiscus gen. nov. (Hyphodiscaceae, Helotiales), with L. epibryus comb. nov. and L. hemiamyloideus sp. nov., two overlooked bryophilous ascomycetes with a wide host range. - Mycol Progress 23, 76. https://doi.org/10.1007/s11557-024-02003-w

Greiff, G. R. L. & Döbbeler, P. (2024) Bryobroma, nom. nov., a replacement for pyrenomycetous bryophilous ascomycetes formerly included in the illegitimate genus Bryomyces Döbbeler. - Ascomycete.org, 16 (2) : 15–16.

Greiff, G. R. L., A. Büschlen & Döbbeler, P. (2024) Studies on Bryocentria brongniartii (Hypocreales, Bionectriaceae) on Frullania dilatata in Europe reveal an anamorph and a cryptic species, B. confusa sp. nov. - Ascomycete.org, 16 (4) : 197–204.

Greiff, G. R. L. & Döbbeler, P. (2024) Endoantria benetecta (Hypocreales) – a new perianthicolous ascomycete on Lejeunea patens in British temperate rainforests. – Nova Hedwigia. 119 (3–4): 351–367.

2023

Döbbeler, P., Davison, P. G. & Buck, W. R. (2023). Bryophilous ascomycetes of North America – an overview of the recorded species. – Herzogia 36: 305 –370.

2022

Janošík, L., Catcheside, P. S., Catcheside, D. E. A. & Döbbeler, P. (2022). Octosporella australis sp. nov. and O. jungermanniarum agg. (Pezizales), two notable hepaticolous fungi from Australia. Nova Hedwigia 115(1-2): 157-179. DOI: 10.1127/nova_hedwigia/2022/0699

2021

Döbbeler, P. & Braun, U. (2021): Pseudomicrodochium bryophilum – a new conidium-forming ascomycete on foliose liverworts. Schlechtendalia 38: 326–330.

Döbbeler P., Büschlen A., Eckstein J. & Gross A.. (2021) Belonium coroniforme Rehm (Helotiales), a highly specialized muscicolous ascomycete on Orthotrichaceae and Leucodon. – Sydowia 74: 163–174. DOI 10.12905/0380.sydowia74-2021-0163

Döbbeler, P. & P. G. Davison 2021. Non-hypocrealean ascomycetes on Frullania mainly from Southeastern North America. – Nova Hedwigia 113(3-4): 361-401. DOI: 10.1127/nova_hedwigia/2021/0663

Döbbeler P., Büschlen A. & Eckstein J. (2021): Octospora bicarpa sp. nov. (Pezizales), the first species of the bryophilous genus Octospora Hedw. with ameroconidia. Ascomycete.org 13(2): 85–91.

Döbbeler P. (2021): Bryomycology without bryology? Nova Hedwigia 112(1–2): 267–270.

2020

Döbbeler P. (2020): Mehltaue, Rost- und Brandpilze der Gurgler Täler (Ötztaler Alpen, Tirol). Mitteilungen des Naturwissenschaftlichen Vereins für Steiermark 150: 7–43.

Brackel W. von & Döbbeler P. (2020): An addition ot the knowledge of lichenicolous fungi of Greece with a key to the lichenicolous fungi on Collema s. l. Folia Cryptogamica Estonica 57: 147–152.

2019

Döbbeler P. & Davison P.G. (2019): Two new leaf-perforating ascomycetes on corticolous liverworts from Southeastern North America. Ascomycete.org 11(6): 222 – 228. (DOI: 10.25664/ART-0278)

Döbbeler P. (2019): Tremella bryonectriae sp. nov. - the first hyperbryophilous fungal species. Nova Hedwigia 109(3-4): 369 – 378.

Sochorová Z., Döbbeler P., Sochor M. & van Rooy, J. (2019): Octospora conidiophora (Pyronemataceae) – a new species from South Africa and the first report of anamorph in bryophilous Pezizales. MycoKeys 54: 49 - 76.

2018

Döbbeler P., Ewald J., Fleischmann A., Meierott, L. & Renner S.S. (2018): Dr. Wolfgang Lippert 1937 - 2018. Erforscher und Schützer der bayerischen Flora. Ber. Bayer. Botan. Ges. 88: 167 – 177.

Döbbeler P., Berger F., Negrín Piñero R. & Vega M. (2018): Octosporella microtricha and O. nematospora – two new pezizalean ascomycetes on Frullania. Herzogia 31: 101 – 108.

Döbbeler P., Müller F. (2018): Bryostroma acrocarpum – a novel muscicolous ascomycete from New Caledonia. Herzogia 31: 109 – 113.

Döbbeler P. (2018): Hypocrealean hyperepiphyllous ascomycetes. Biosyst. Ecol. Ser. 34: 187 – 225. Wien, Verlag der Österr. Akad. Wiss.

Döbbeler P., Linde J. & E. Rubio (2018): Bryocentria octosporelloides (Hypocreales) – a new species on Cololejeunea minutissima from Asturias (Spain). Ascomycete.org 10(2): 77 – 80.

Huhtinen, S. & P. Döbbeler (2018): Bryoscyphus hyalotectus (Helotiales), a new polytrichicolous ascomycete from North America. Karstenia 57 (2017): 33 – 36.

2017

Döbbeler P. & Buck W.R. (2017): Dactylospora inopina (Lecanorales), a new biotrophic parasite on Radula (Hepaticae) from the Cape Horn Archipelago, Chile. Nova Hedwigia 105(1-2): 87 – 93.

Döbbeler P. (2017): Antheridia of Bryum capillare release moving spermatozoids after two months in the air-dry state. Herzogia 30(2): 501 – 503.

Döbbeler P. & P.G. Davison (2017): Frullania as a hotspot for hypocrealean ascomycetes: ten new species from Southeastern North America. Nova Hedwigia 106(1-2): 209 – 256.

Egertová Z., Döbbeler P. & M. Sochor (2017): Octosporopsis erinacea and Octospora kelabitiana (Pezizales) – two new hepaticolous ascomycetes from Borneo. Mycological Progress 17: 103. (DOI: 10.1007/s11557-017-1354-5

2016

Döbbeler P. (2016): Three new ascomycetes on epiphyllous liverworts. Karstenia 56: 47 – 53.

2015

Nordén B., Gardiennet A., Priou J.-P. & Döbbeler P. (2015): Bryocentria hypothallina (Hypocreales) – a new species on Metzgeria furcata. - Ascomycete.org. 7(4): 121 – 124.

Döbbeler P., Davison P.G. & Buck W.R. (2015): Two new hypocrealean ascomycetes on bryophytes from North America. Nova Hedwigia 100: 383 – 390.

2014

Döbbeler P. & Facher E. (2014): Octospora mnii (Pezizales), a new ascomycete on the persistent protonema of Rhizomnium punctatum. - Karstenia 54: 49 – 56.

2013

Döbbeler P. & Hertel H. (2013): Bryophilous ascomycetes everywhere: Distribution maps of selected species on liverworts, mosses and Polytrichaceae. – Herzogia 26: 361 – 404.

2012

Döbbeler P. (2012): Gynonectria intraspora gen. et sp. nov. (Hypocreales), a perianthicolous phyllosphere fungus with a novel ascospore type. – Mycological Progress 11: 473 – 481.

2011

Döbbeler P. (2011): Two new hepaticolous species of the genus Octosporella (Pezizales). – Herzogia 24: 357 – 365.

Döbbeler P. (2011): Bryonectria protonematis (Hypocreales) on Ephemeropsis tjibodensis, the first muscicolous ascomycete of the phyllosphere. – Nova Hedwigia 92: 369 – 376.

2010

Marsh T., Döbbeler P., Huhtinen S. & Stenroos S. (2010): Ascomycetes and anamorphic fungi growing on Plagiochila (Hepaticae) in Finland. – Karstenia 50: 59 – 72.

Döbbeler P. (2010): New species and records of Bryocentria – a hypocrealean genus of bryophilous ascomycetes. – Karstenia 50: 11 – 23.

Stenroos S., Laukka T., Huhtinen S., Döbbeler P., Myllys L., Syrjänen K. & Hyvönen J. (2010): Multiple origins of symbioses between ascomycetes and bryophytes suggested by a five-gene phylogeny. – Cladistics 26: 281 – 300.

Huhtinen S., Laukka T., Döbbeler P. & Stenroos S. (2010): Six novelties to European bryosymbiotic discomycetes. – Nova Hedwigia 90: 413 – 431.

Döbbeler P. (2010): Hyperepiphyllous, perianthicolous Hypocreales – highly specialized ascomycetes of the phyllosphere. – Mycologia 102: 404 – 417. (Abstract)

2008

Döbbeler P. (2008): Ascomycetes on Pogonatum urnigerum (Musci). – Nova Hedwigia 87: 129 – 144.

2007

Döbbeler P. (2007): Ascomycetes on Polytrichadelphus aristatus (Musci). – Mycological Research 111: 1406 – 1421.

Eckstein J. & Döbbeler P. (2007): Fungal spores within sporophytes of Gymnostomum viridulum. – Journal of Bryology 29: 169 – 173.

2006

Döbbeler P. (2006): Ascomycetes on Frullania dilatata (Hepaticae) from Tuscany. – Mycological Progress 5: 32 – 40.

Döbbeler P. (2006): Two new polytrichicolous Ascomycetes from Ecuador. – Nova Hedwigia 82 (1 – 2): 257 – 264.

2005

Döbbeler P. (2005): Three new hypocrealean Ascomycetes on bryophytes. – Sydowia 57 (2): 179 – 188.

Döbbeler P. (2005): Ascospore diversity of bryophilous Hypocreales and two new hepaticolous Nectria species. – Mycologia 97: 924 – 934.

2004

Döbbeler P. (2004): Bryocentria (Hypocreales), a new genus of bryophilous Ascomycetes. – Mycological Progress 3: 247 – 256.

Döbbeler P. (2004): Octosporella erythrostigma (Pezizales) and Pithyella frullaniae (Helotiales), two remarkable ascomycetes on Frullania dilatata. – Feddes Repertorium 115: 5 – 14.

Döbbeler P. & Feuerer T. (2004): Stenocybe nitida (Mycocaliciales), an unusual ascomycete on Plagiochila punctata. – Biblioth. Lichenol. 88: 91 – 102.

Döbbeler P. & Rambold G. eds. (2004): Contributions to Lichenology. Festschrift in Honour of Hannes Hertel. Biblioth. Lichenol. 88: 1 – 739.

2003

Döbbeler P. (2003): Ascomycetes on Dendroligotrichum (Musci). – Nova Hedwigia 76: 1 – 44.

Lippert W. & Döbbeler P. (2003): Dr. Helmut Wunder 30.1.1940 – 17.12.2001. – Ber. Bayer. Bot. Ges. 72 (2002): 201 – 203.

2002

Döbbeler P. (2002): Microniches occupied by bryophilous ascomycetes. – Nova Hedwigia 75: 275 – 306.

2001

Döbbeler P. (2001): Polytrichadelphus magellanicus sensu lato (Musci) and its ascomycetes – different fungi on different hosts. – Sendtnera 7: 35 – 45.

Döbbeler P. & Triebel D. (2001): Potridiscus polymorphus (Leotiales) – a new ascomycete on Polytrichaceae (Musci) with palaeoaustral distribution. – Hoppea 61 (2000): 71 – 83.

1999

Döbbeler P. (1999): Two new species of Bryonectria (Hypocreales, Ascomycetes) on bryophytes. – Sendtnera 6: 93 – 102.

Döbbeler P. (1999): Polytrichadelphus magellanicus – a mycological Eldorado: five new ascomycetes on a single collection from Tierra del Fuego. – In: Anonymus (ed.), Riclef-Grolle-Festschrift. – Haussknechtia Beih. 9: 79 – 96. Jena.

Döbbeler P. (1999): Lamellen- und Epidermishaare als neues Merkmal der Polytrichaceen (Musci). – Nova Hedwigia 69: 357 – 380.

1998

Döbbeler P. (1998): Ascomyceten auf der epiphyllen Radula flaccida (Hepaticae). – Nova Hedwigia 66: 325 – 373.

Döbbeler P. (1998): Epibryon pedinophylli – a new leaf-perforating ascomycete on Pedinophyllum interruptum (Hepaticae). – Sendtnera 5: 19 – 27.

1997

Döbbeler P. (1997): Biodiversity of bryophilous ascomycetes. – Biodiversity and Conservation 6: 721 – 738.

1996

Döbbeler P. (1996): Gloeopeziza cuneiformis (Leotiales, Ascomycetes) – ein neuer Lamellenbewohner von Polytrichum. – Sendtnera 3: 103 – 109.

Döbbeler P. (1996): Potriphila navicularis gen. et sp. nov. (Ostropales, Ascomycetes), ein bipolar verbreiteter Parasit von Polytrichum alpinum. – Nova Hedwigia 62: 61 – 77.

1995

Döbbeler P. (1995): Myxophora tjibodensis spec. nov. (Dothideales), ein weiterer Symbiont der Blaualge Scytonema. – In: Farkas, E. É., Lücking, R. & Wirth, V. (eds), Scripta lichenologica – lichenological papers dedicated to Antonin Vezda. – Biblioth. Lichenol. 58: 73 – 80.

Döbbeler P. (1995): Burenia myrrhidendri spec. nov. (Protomycetales), ein bemerkenswerter biotropher Ascomycet in den Früchten einer baumförmigen Umbellifere aus Costa Rica. – Nova Hedwigia 60: 171 – 177.

1994

Döbbeler P. (1994): Epigloea urosperma (Ascomycetes) – ein neuer Flechtenparasit. – Sendtnera 2: 277 – 282.

1993

Döbbeler P. (1993): Hifas intracelulares en hepáticas y musgos como posible fuente de error. – Briolatina 31: 2 – 5.

Döbbeler P. (1993): Nectria sanramonensis (Hypocreales, Ascomycetes), un nuevo parásito briófilo de Costa Rica. – Brenesia 37 (1992): 145 – 149.

Döbbeler P. (1993): The infecting structures of Octospora roxheimii (Pezizales). – Arnoldia 6: 12 – 14.

Döbbeler P. & Carranza J. (1993): Cuatro especies nuevas de Nectria (Hypocreales: Ascomycetes) en hepáticas epífilas de Costa Rica. – Rev. Biol. Trop. 41: 203 – 208.

1992

Döbbeler P. & Menjívar R. (1992): Tres nuevas especies de ascomicetes en hepáticas epífilas de Costa Rica. – Rev. Biol. Trop. 40: 73 – 81.

1988

Döbbeler P. (1988): Drei neue moosbewohnende Ascomyceten. – Pl. Syst. Evol. 158: 329 – 340.

1987

Döbbeler P. (1987): Ascomycetes growing on Polytrichum sexangulare. – In: Laursen, G. A., Ammirati, J. F. & Redhead, S. A. (eds), Arctic and alpine mycology II. – Environm. Sci. Res. 34: 87 – 107.

1986

Döbbeler P. (1986): Ascomyceten auf Polytrichaceen. – Sydowia 38 (1985): 41 – 64.

Döbbeler P. (1986): Belonioscyphella hypnorum (Helotiales, Ascomycetes), ein nekrotropher Parasit auf Laubmoosen. – Ber. Bayer. Bot. Ges. 57: 153 – 158.

Döbbeler P. (1986): Ein Zweitfund von Epicoccum plagiochilae (Hyphomycetes). – Mitt. Bot. Staatssamml. München 22: 485 – 492.

Döbbeler P., Poelt J. & Vezda A. (1986): Lopadium hepaticola spec. nov., ein moosparasitisches echtes Lopadium von der Südhalbkugel. – Herzogia 7 (1985): 81 – 91.

1985

Döbbeler P. & Triebel D. (1985): Hepaticole Vertreter der Gattungen Muellerella und Dactylospora (Ascomycetes). – Bot. Jahrb. Syst. 107: 503 – 519.

Döbbeler P. (1985): Moosbewohnende Ascomyceten VII. Neufunde einiger Arten der Gattung Epibryon. – Mitt. Bot. Staatssamml. München 21: 757 – 773.

Döbbeler P. (1985): Nachruf: Heinz Itzerott (1912 – 1983). – Z. Mykol. 51: 165 – 166.

1984

Döbbeler P. (1984): Symbiosen zwischen Gallertalgen und Gallertpilzen der Gattung Epigloea (Ascomycetes). – In: Hertel, H. & Oberwinkler, F. (Herausg.), Festschrift J. Poelt. – Beih. Nova Hedwigia 79: 203 – 239.

Döbbeler P. (1984): Einige für Bayern neue bryophile Ascomyceten. – Ber. Bayer. Bot. Ges. 55: 79 – 84.

Döbbeler P. & Hertel H. (1984): Drei neue moosbewohnende Ascomyceten aus der Subantarktis (Marion Island). – Sydowia 36 (1983): 33 – 45.

1983

Döbbeler P. (1983): Hypobryon gen. nov. (Dothideales), eine bemerkenswerte Gattung bryophiler Pyrenomyceten. – Nova Hedwigia 37: 1 – 17.

Döbbeler P. & Itzerott H. (1983): Ein Neufund des muscicolen Chytridiomyceten Pleotrachelus wildemanii. – Mitt. Bot. Staatssamml. München 19: 431 – 434.

1982

Döbbeler P. & Vezda A. (1982): Macentina hepaticola, eine neue Flechte aus Zaire. Mitt. Bot. Staatssamml. München 18: 1 – 7.

Döbbeler P. (1982): Moosbewohnende Ascomyceten VI. Einige neue Pyrenomyceten. – Mitt. Bot. Staatssamml. München 18: 341 – 358.

Döbbeler P. & Futschig J. (1982): Über Fremdsporen in den Kapseln von Anthoceros. – Herzogia 6: 271 – 282.

Itzerott H. & Döbbeler P. (1982): Octospora meslinii und O. rubens (Pezizales), zwei weitere bryophile Gallbildner. – Mitt. Bot. Staatssamml. München 18: 201 – 211.

1981

Döbbeler P. (1981): Moosbewohnende Ascomyceten V. Die auf Dawsonia vorkommenden Arten der Botanischen Staatssammlung München. – Mitt. Bot. Staatssamml. München 17: 393 – 473.

Döbbeler P. (1981): Phoma plagiochilae, ein interzellulärer lebermoosbewohnender Pyknidienbildner. – Mitt. Bot. Staatssamml. München 17: 231 – 236.

Döbbeler P. & Itzerott H. (1981): Zur Biologie von Octospora libussae und O. humosa, zwei im Moosprotonema wachsenden Pezizales. – Nova Hedwigia 34: 127 – 136.

Döbbeler P. & Poelt J. (1981): Arthopyrenia endobrya, spec. nov., eine hepaticole Flechte mit intrazellulärem Thallus aus Brasilien. – Pl. Syst. Evol. 138: 275 – 281.

1980

Döbbeler P. (1980): Moosbewohnende Ascomyceten IV. Zwei neue Arten der Gattung Octosporella (Pezizales). – Mitt. Bot. Staatssamml. München 16: 471 – 484.

Döbbeler P. (1980): Phycorella scytonematis (Dothideales), ein neuer Symbiont der Blaualge Scytonema. – Sydowia 33: 33 – 38.

Döbbeler P. (1980): Epibryon endocarpum sp. nov. (Dothideales), ein hepaticoler Ascomycet mit intrazellulären Fruchtkörpern. – Z. Mykol. 46: 209 – 216.

Döbbeler P. (1980): Untersuchungen an moosparasitischen Pezizales aus der Verwandtschaft von Octospora. – Nova Hedwigia 31: 817 – 864.

1979

Döbbeler P. (1979): Moosbewohnende Ascomyceten III. Einige neue Arten der Gattungen Nectria, Epibryon und Punctillum. – Mitt. Bot. Staatssamml. München 15: 193 – 221.

Döbbeler P. (1979): Vezdaea dawsoniae (Lecanorales), ein neuer Moosbewohner aus Neuguinea. – Herzogia 5: 95 – 101.

Döbbeler P. (1979): Taphrina bergeniae, ein neuer parasitischer Ascomycet aus dem Botanischen Garten in München. – Sydowia 31 (1978): 30 – 37.

Döbbeler P. (1979): Moosbewohnende Ascomyceten II. Acrospermum adeanum. – Mitt. Bot. Staatssamml. München 15: 175 – 191.

Döbbeler P. & Nannenga-Bremekamp N.E. (1979): Licea gloeoderma, ein neuer Myxomycet aus Bayern. – Z. Mykol. 45: 235 – 238.

Poelt J. & Döbbeler P. (1979): Bryostigma leucodontis nov. gen. et spec., eine neue Flechte mit fast unsichtbaren Fruchtkörpern. – Pl. Syst. Evol. 131: 211 – 216.

1978

Döbbeler P. (1978): Moosbewohnende Ascomyceten I. Die pyrenocarpen, den Gametophyten besiedelnden Arten. – Mitt. Bot. Staatssamml. München 14: 1 – 360.

1977

Döbbeler P. & Poelt J. (1977): Absconditella celata spec. nov., eine Flechtenart aus Lappland. – Herzogia 4: 363 – 366.

Poelt J. & Döbbeler P. (1977): The genus Vezdaea: a supplement. – Lichenologist 9: 169 – 170.

1976

Döbbeler P. & Remler P. (1976): Über einige neue oder bemerkenswerte Myxomyceten der Steiermark. – Mitt. Naturwiss. Vereins Steiermark 106: 131 – 141.

1975

Poelt J. & Döbbeler P. (1975): Lecidea margaritella, eine an ein Lebermoos gebundene Flechte, und ihr Vorkommen in Mitteleuropa. – Herzogia 3: 327 – 333.

Poelt J. & Döbbeler P. (1975): Über moosparasitische Arten der Flechtengattungen Micarea und Vezdaea. – Bot. Jahrb. Syst. 96: 328 – 352.

1974

Döbbeler P. & Poelt J. (1974): Beiträge zur Kenntnis moosbewohnender Discomyceten. I. Die Gattung Bryodiscus. – Svensk Bot. Tidskr. 68: 369 – 376.

Research interests

Taxonomy and biology of bryophilous ascomycetes with emphasis on Central European species, and species infecting epiphyllous hepatics and Polytrichaceae

Ascomycetes on bryophytes: systematics and biology

Bryophilous ascomycetes are ecologically defined as lichenicolous, coprophilous or marine fungi. The only common character they share is their occurrence on bryophytes. Hepaticolous fungi infect both thallose and foliose hepatics, while muscicolous species infect mosses. More specific terms are sometimes used for the parasites of some well-known and frequently infected taxa, e.g. frullaniicolous species occur on Frullania, sphagnicolous species on Sphagnum, and polytrichicolous species on Polytrichaceae.

Perianthicolous species attack the developing sporophytes within the perianths of hepatics. Bryophily is a life strategy that has evolved independently in many non-related fungal lineages. Accordingly, bryophilous species represent a heterogeneous assemblage of symbionts of quite different systematic position, mode of nutrition, host selection and geographical distribution. Due to this exceptionally high diversity bryophilous ascomycetes exhibit many characters that are known in other ascomycetes. Bryomycology is the study of these fungi. It provides a plethora of novel fungal taxa, unrecorded hosts, and hitherto unknown characters, such as the formation of intracellular ascomata, leaf-perforating ascomata, unique ascospore types, and subterranean rhizoid galls.